Graptolites were colonial animals
that lived from Upper Cambrian through Lower Carboniferous
(Mississippian) times. Since the use of electron microscopy was introduced to
their investigation in the 1970s, they have been considered to be allied with
the pterobranchs, a rare group of modern marine animals belonging in the phylum
Hemichordata. Their name was derived from a combination of two Greek words, graptos,
meaning "written", and lithos, meaning "rock", because
Linnaeus, who described them first, thought of them as images resembling
fossils rather than being true fossils. Each graptolite colony has a variable
number of branches originating from an initial individual, with each subsequent
individual (or zooid) being housed within a tubular or cup-like structure. These
fossils are rare in Devonian rocks of south central Oklahoma because they do not survive surface exposure well,
and it is likely that grazers and scavengers cleaned up their dead debris.
Most of the dendritic or
many-branched types, such as those shown here, are classified as dendroid
graptolites (order Dendroidea). They appeared in the Cambrian period and were
generally attached to the sea-floor by a root-like base. Graptolites with
relatively few branches were derived from the dendroid graptolites at the
beginning of the Ordovician period; and colonies of this latter group
(order Graptoloidea) were pelagic, drifting freely on the sea surface or
attached to floating seaweed. The Graptoloidea were successful and
prolific, being important members of the plankton until they died out in the early Devonian period.
The dendroid graptolites survived into Lower Carboniferous times.
Their
worldwide distribution, rapid evolution and good preservation have allowed graptolite
use as index fossils and for division of Ordovician and Silurian rocks into
distinct graptolite zones, each less than one million years in duration. According to the entry on Wikipedia, from
which I liberally borrowed the information here, graptolites have been used to
estimate water depth and temperature. I will follow up on this; as such a
correlation may help further refine the conditions in which the rocks of Black Cat Mountain
were deposited. |